The journal publishes topics across the life sciences. As long as the core subject lies within the biological sciences, some issues may also include content crossing into other areas such as the physical sciences, social sciences, biophysics, policy, economics etc. Issues generally sit within four broad areas (although many issues sit across these areas):
Organismal, environmental and evolutionary biology
Neuroscience and cognition
Cellular, molecular and developmental biology
Health and disease.
SCI EI SSCI SCOPUS